Grumpy Cat, the feline famous on the internet for her permanent scowl, has died aged seven, her owners say.
A statement says she died on Tuesday following complications from a recent urinary tract infection.
The cat from Arizona had "helped millions of people smile".
Grumpy, whose real name was Tardar Sauce, went viral in 2012 after photographs of her sour expression emerged online. Her image quickly spread as a meme.
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